Norway Spruce - Picea abies

 

 

On this page we present a chapter on Norway Spruce, reproduced from "British Forest Trees and their Sylvicultural Characteristics and Treatment"  by John Nisbet, published in 1893.
As you might expect from a book with such a title, Nisbet writes predominantly from the point of view of a forester interested in how to grow the tree and look after it. You will find information on other aspects of this tree elsewhere on our website (see below).
The essay may be well over 100 year old, but it contains a wealth of information about the Norway Spruce, which you won't easily find in more modern writing. 

 

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Norway Spruce - Picea abies

Sylviculture of Norway Spruce, Picea Abies

 

BRITISH FOREST TREES

AND THEIR

SYLVICULTURAL CHARACTERISTICS

AND TREATMENT

BY

JOHN NISBET, D.OEc.

OF THE INDIAN FOREST SERVICE;

TRANSLATOR OP FÜRST'S  "PROTECTION OF WOODLANDS."

 

 

 

 

London

MACMILLAN AND Co.

 AND NEW YORK

1893

The Right of Translation and Reproduction is Reserved

 


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Our reproduction starts from page 83 from the section:

"CHIEF SPECIES, forming, or capable of forming, Pure Forests."

"Conifers"

and includes all pages up to page 114


SPRUCE, NORWAY SPRUCE, OR SPRUCE FIR (Pinus abies, L =  Pinus picea, Du Roi = Pinus  excelsa, Lam. = Abies excelssa, D.C.  =  PICEA EXCELSA, Link).

Distribution. ---In this respect the spruce is inferior only to the Scots pine. It extends from latitude 690 N. throughout the whole of northern and central Europe, southwards to the slopes of the Alps, Cevennes and Pyrenees. It forms extensive forests in Scandinavia, Finland, Lapland, and Russia, although its growth there is not to be compared with that attained on the hilly land and mountain masses in central Germany. It seems not to have been indigenous to Scotland or England, for no fossil traces of it have been found, and no historical record exists of it having ever formed forests on the hills of ancient Britain; it was probably only introduced toward the middle of the sixteenth century.
In France, as well as in Scotland on an altogether smaller scale, it has been cultivated to a much less extent than other conifers, and in Spain, Italy and Greece it is seldom met with forming forests. The eastern limit of the species is not easy to fix, as it gradually merges into another variety, the Siberian spruce (Picea obovata).
It ascends the Harz mountains to about 3,300 feet, the Black Forest and the mountains of Silesia to 3500---4000 feet, the Bavarian Alps to nearly 6,000 feet, and the central Alpine ranges to over 6600 feet.  In Germany and Switzerland the spruce is the principal forest tree on all mountain ranges and hilly tracts, often forming pure forests over large tracts of country, whilst below it there is a girdle of deciduous broad-leaved trees into whose domain it is always trying to extend its frontiers.
On the lower hills it is frequently found mixed with the silver fir and the beech, and at higher elevations in lower latitudes with the larch; in Silesia and East Prussia it forms extensive forests, often in company with the Scots pine. It is emphatically a tree of the upper hilly region, the more so in proportion as the hills and mountains are massive in formation, instead of consisting of a series of ridges and chains; it ascends to its greatest height when the general elevation of the surrounding country is considerably above the sea-level. Where the uplands fall away towards warm, dry, lowland tracts the spruce is not indigenous, and though pure forests of it are frequently to be found on indifferent soils in these localities, they are almost always the result of artificial production, and too often but poor in their production of timber.

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Tree-form and Roof-system of the spruce exhibit many differences from, and indeed direct contrasts to, those of the Scots pine. Its roots are mostly confined to the upper layer of the soil, and these of one tree often interlace with those of its nearest neighbours, thus obtaining some little protection during high winds. Its horizontal root-system, seldom going lower than 18-20 inches, and unprovided with any deep-reaching tap-root, stamps it indeed as the tree of shallow-soiled mountains, but offers it too often a sacrifice to the winds. It develops great numbers of rootlets, and as it also possesses the property of extending its roots to a great distance, it has a comparatively large area from which to draw supplies of nutriment, although owing to its density when forming pure forests the individual growing-space is limited beyond that of any other tree except perhaps the silver fir.
Although slower in growth during youth than the Scots pine, it maintains a much steadier rate of increase in height and grows up in dense canopy without much tendency to branch formation or interruption. In height, length of bole, straightness and full-woodedness of stem, freedom from branches, greatest quantitative production of wood and of useful timber per acre, it is rivalled only by the silver fir. Its crown is cone-shaped, and when grown in isolated positions the whole foliage is retained in more or less conical form from the summit downwards to near the ground; the short leaves or needles on the branches are retained for five to seven years.

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Requirements as to Soil and Situation. -----Shallow-rooted though the spruce undoubtedly be, yet it demands freshness in the soil, and cool, damp, mountain atmosphere is beneficial to its growth. Where other woods shelter it from the wind it also finds a suitable abode in the vicinity of the seacoast, but its best development is attained in protected localities on mountain sides. In its true home the average temperature in July does not much exceed 660° Fahr. (Willkomm, Die forstliche Flora Deutschlands und Oesterreichs, 887, p. 81.) and the total minimum warmth requisite during each annual period of active vegetation has been ascertained to be about 2,610° Fahr., which is about the annual average quantity of warmth developed at latitude 69° north. Its growth seems best when the enjoyment of the warmth, and thereby the period of active vegetation, is confined to a short summer season, during which there is daylight for the longest possible
time, as in the north of Scotland and in Norway and Sweden. Drought is less easily borne by the spruce than by any other tree.
In regard to the quality of the soil, spruce shows a considerable degree of indifference, or at any rate adaptability; it makes greater demands on mineral strength than the Scots pine, but is content with less than the silver fir. Being at the same time one of the thickly-foliaged species of trees that improve the soil, and recruit such soils as have become impoverished, it is frequently found where broad leaved deciduous trees have previously allowed the soil to become deteriorated and impoverished.
Spruce thrives on soils of the most varied description, from the strong, friable mountain soil down through the binding varieties to the sandy-loamy, and the drained moors and bogs. Soils that show any good growth of weeds like Epilobium, Senecia, Atropa, or Digitalis, or of grasses like Carex are generally fresh and capable of producing good spruce forests, as also are those with high growth of whortleberry; but on tracts covered with heath and heather plantations should more frequently be made with spruce and Scots pine, than with spruce alone. It is not indifferent to mineral strength, but the chief factor in determining the suitability or non-suitability of any particular locality is certainly the equable distribution of a moderate quantity of moisture throughout the soil. The older sand formations, and loamy deposits resting on limy subsoil found in Alpine districts bear good spruce forests, but on limy soils it is apt to suffer from fungous diseases. On marls, loams, and rich clayey soils, deciduous broad-leaved trees find a more suitable home than the spruce. Low sandy plains with dry gravelly soil, soured undrained stretches with stagnant soil-moisture, moorpan with excess of sesquioxide of iron, or tracts liable to inundation are not the localities on which spruce can be expected to attain its normal development.
The most suitable aspect depends on the locality, and the elevation above the sea-level. Towards the lower limit of its proper region it prefers the cool, moist, north and north-east exposures, whilst towards the upper limit it seeks the southern and south-western aspects in order to obtain the requisite degree of warmth, and to escape from the drying-up influence of the east winds.

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Requirements as to Light.---The ability to retain its branches in foliage for five to seven years, and the consequent density of its crown, give indication of the large capacity with which the spruce is endowed as regards bearing shade; but the extent to which the demand for some measure of enjoyment of light exists, is mainly dependent on how far any particular locality varies from the normal situations suitable to it.
Where these various climatic changes are distinctly discernible, the demand for light becomes greater, and the capacity for bearing shade smaller. Where soils are wanting in moisture, young spruce cannot thrive under standards which intercept and partially retain the atmospheric precipitations. Excess of light on the other hand stimulates to increased assimilation of sap and too rapid growth in the earlier stages, which in consequence seriously affects the quality of the timber produced. In its true home, as, for example, on the fresh loamy soil of the Bavarian plateau, it frequently has to content itself for the first fifteen to twenty years of its existence with only a moderate supply of light under scattered standard parent trees, before being gradually admitted to the full enjoyment of unrestricted light and sunshine and the opportunity of normal development.

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Attainment of Maturity and Reproductive Capacity.--- Spruce is generally grown with a rotation of seventy to eighty years for ordinary timber, or one hundred to one hundred and twenty years for the production of larger assortments, but local market considerations must determine when the fall can most advantageously take place; higher periods of rotation are only advisable where the quality of the soil is above the average. Good money returns on the capital represented point decidedly towards the growth of spruce (and Douglas Fir) as being one of the most remunerative and profitable ways of utilising forest soils of about average quality.
Good seed years are less frequent with the spruce than the Scots pine, but are generally - reckoning from the fiftieth to sixtieth year - more productive when they occur; on the Harz mountains one good and one minor seed year are expected in every six years. Seed years can be foretold by the flower-buds, and the twigs showing these, broken off by squirrels and birds which feed on them. The ruddy-brown seed ripens in the October after the flowering, and is scattered from the cones in spring; it is somewhat larger and heavier than the greenish-black or brownish seed of Scots
pine, and is reckoned good in quality when test experiments show a germinative power of seventy-five to eighty per cent. lit thus has not only a greater germinative power than the Scots pine, but it retains this somewhat longer, especially when kept in the cone. Each cone has 200 to 250 seeds, and one pound contains from 55,000 to 60,000 seeds.

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Liability to suffer from External Dangers.---As an offset against its many excellent qualities, spruce has unfortunately to contend with many external dangers at all periods of its existence, here of course to a greater extent, and there to a less, according to the soil and situation. Sharp, biting winds hinder reproduction at high elevations, except under the shelter of protective standards. Frost is only liable to damage the young growth at its earliest stage. Accumulations of snow, and of ice formed after rain on the heavy foliaged branches, bend down saplings in thickets, break the poles in young forests, snap off the tops of trees, and make large holes here and there in the canopy, especially at moderate elevations (on the Harz mountains, particularly those between 1700 - 2300 feet) where the snow is larger in flake than at high altitudes. Dense forests suffer most from snow, whilst a larger growing-space increases the danger from hanging ice.
No other species of forest tree is less able than the spruce to resist the violence of storms. Its shallow root-system, the long lever formed by the bole, and the purchase obtainable by the wind on the dense crown of foliage near the summit, all combine to weaken the resistance it is able to offer, especially in early spring and late autumn when strong winds are frequent just at the time when the foliage is often heavy with moisture and the soil sodden and softened by continuous rainfall. Whole forests are then often thrown down. The danger from wind is considerably lessened when other species are grown in admixture with it, or when natural reproduction takes place by the annual or periodical removal of the largest trees only.
Drought is injurious in the youngest stages of growth especially when accompanied by dry winds.
On unsuitable localities the mature stems are somewhat liable to die off, and old tree forests often suffer from fungous diseases, occasioned chiefly by Trarnetes pini and Nectria cucurbitula on the stem and branches, and by Trametes radiciperda and Agaricus melleus in and near the roots, which diminish the value of the timber. Cotyledons and leaves of seedlings are demolished by Phytophthora omnivora. In young plantations, and particularly in nurseries and young seedling crops, blanks are often caused after wet summers by Pestalozzia Hartigii. Red-rot in the timber is occasioned by Polyporus vaporarius, and white-rot by Polyporus borealis or, less frequently, P. fulvus.
But even the climatic dangers and fungous diseases combined are surpassed in importance by those to which spruce is exposed at all periods of its growth and development from the attacks of insect enemies. Extensive tracts of pure spruce forest have recently in Germany been severely damaged, partially destroyed, and even often killed outright by insects, the lower elevations suffering far more severely than those situated within the true mountainous region.
To enumerate merely the more important of such enemies, larvæ of Melolontha vulgaris and hippocastani, and Gryllotalpa vulgaris, and the full-grown Hylastes cunicularis and Gryllotalpa do great damage to the roots of seedlings and young plants, whilst the beetles Hylobius abietis and
pinastri and Hylastes cunicularis gnaw the tender bark of their stems; later on the cortex and sap-wood suffer from both the larvæ and the fully-developed beetle of three varieties of Bostrychus  - amitinus, calcographus, and typographus, whilst the mature wood is damaged by both the active forms of Xyloterus lineatus. Buds and foliage are destroyed completely, and valuable forests utterly ruined over immense tracts of country, by that scourge of the coniferous forests of Germany, Liparis monacha, the voracity of whose caterpillars is only equalled by their almost unlimited numbers in bad years like 1889, 1890, 1891 in southern Bavaria, where alone the extent of forests devastated by the black arches, "nun," or spruce moth (Liparis monacka) is estimated to be about 42,500 acres or sixty-six square miles, of which by far the greatest portion was pure forest of spruce.
(For a detailed account of this insect and its ravages, see the Transactions of the Highland and Agricultural Society for 1893.)  In the mixed forests attacked, the beech and Scots pine suffered comparatively much less than the spruce, although they were also badly injured; the spruce, however, was usually killed outright, owing to the much smaller reserves of starchy matters stored up by this species for subsequent constructive purposes.
In recuperative power with regard to injuries received, whether caused by insects or by deer, the thin-barked spruce is not well endowed. Where a strong head of game is maintained, red-deer do more damage in spruce forests than elsewhere, by stripping the bark with their teeth during summer, as well as for food during winter. The damage caused is often very serious, and is generally most widespread in pole-forests from twenty to forty years of age, although it is often done also in tree-forests up to sixty years of age. For the healing of the wounds thus caused, spruce has unfortunately less recuperative power than the silver fir or even the Scots pine.

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Sylvicultural Treatment of Spruce.---The economic value of spruce is by no means small, if its cultivation takes place under suitable circumstances. Moderate in its demands on soil, which it also protects and improves in quality, and seldom giving much trouble in the formation and reproduction of forests, spruce yields on favourable localities a larger outturn of timber than any other tree usually grown in pure-forests (vide table on page 44), without requiring a high period of rotation to attain marketable proportions. It remains long in close canopy, and forms lofty, cylindrical, straight stems, that are valuable and of great general utility. It also yields fair returns in the way of thinnings, and occasionally affords good grazing for cattle. From the actuarial point of view many advantages point towards the cultivation of spruce as one of the most remunerative forms of high forest; but the relatively high returns promised can only be realised on soils and situations which admit of the normal development of this species, and these are to be found chiefly in sheltered localities of mountainous regions haying a moist and moderately good soil.
Misled by tempting actuarial calculations, it would be a mistake to transform existing crops into spruce woods without other definite reasons, for most other trees, and particularly the broad-leaved deciduous species, have undoubted advantages over spruce in respect to the greater security they afford against destruction of the crop from snow, storms, or insects. But in mountainous tracts, on undulating soil temporarily reduced by too open crops of deciduous trees, or where timber prices are good, but fuel is little in demand, spruce forests generally as a matter of fact yield the most remunerative returns on soils of about the average quality.
Young crops of spruce do not form canopy so soon as those of Scots pine, though in plantations it is attained earlier than by sowing or with natural reproduction. On sunny localities the soil is apt to become overgrown with weeds like Atropa, Digitalis, Epilobium, Rubus and Urtica, with Vaccinium, Carex, Scirpus and Juncus on the moister patches, which usually interfere with the growth of the young plants and occasionally choke them altogether; but on dry and shallow soil the dangers arising from dry winds and direct insolation are greater. Frost in general does more damage, by lifting the young plants out of the ground, than is occasioned by the actual degree of cold to which they are exposed. Young growth, particularly in woods that have been formed by planting, is greatly exposed to danger from the large brown pine weevil (Hylobius abietis), which bores into the young shoots, and often ruins whole crops formed where the stumps of the mature crop have neither been grubbed up nor barked after the harvesting of the mature fall of timber.
Until the young crop forms canopy, the growth of the individual plants is rather towards lateral extension than in the direction of increase in height, but when once they have closed up (which usually occurs about the twelfth or twentieth year according to the quality of the soil), and the twig-shoots begin to interlace, the development of the leading shoot becomes vigorous, the more so in proportion to the density of the crop and the quality of the soil. As they are densely foliaged and make little demand on growing-space, young thickets of spruce generally stand very thick, and completely cover the soil. So much so is this the case, that, where sowings have been too thick, the development is greatly interfered with; but in general the natural selection of the predominating stems, to form the future crop, goes on normally and quickly, the current annual increase in height culminating with shoots averaging one and a half feet on soils of the best quality between the twenty-sixth and fortieth year, and later, with of course smaller averages, on those of merely average or inferior quality (vide tables on page 36 - 38). Throughout this period of energetic development, the natural suppression of dominated stems gradually progresses, but without any practical interruption of the canopy taking place, so that the boles are enabled to assume that full-wooded cylindrical shape which renders them so valuable. According to Baur this maximum of approach to the cylindrical is attained when the average height of the crop is from sixty-six to eighty feet, but with advancing age it sinks only gradually.
One decided drawback of the close canopy and even development of spruce woods at this stage of their life-history is the consequent danger from accumulations of snow, which often occasion serious damage---a danger however not so much to be feared in either Scotland or England as on the Continent with its severe winters. Later on the density of the crop also gradually diminishes without the continuity of the canopy being very seriously interrupted, whilst from the time that it has reached the tree-forest stage of development, a rich growth of mosses (Hypnum) covers the soil, which, however, gives place to whortleberry (Vaccinium) and similar weeds when self-thinning with consequent interruption of canopy has gone too far. It is at this stage of growth that pure forests of spruce of equal age are most exposed to the danger of windfall and to attacks of bark-beetles (Bostrichini). No species is so little able to resist the force of storms as the spruce, and when once violent winds succeed in breaking up the canopy, they seem to act in a concentrated and cyclonic manner, throwing down everything that offers resistance to their passage.  According to Burckhardt
(Säen und Pftanzen, 1880, p. 329. The introduction of the silver fir and the re-introduction of beech into these spruce forests has been occupying the attention of foresters there for many years past.)  in the Hanoverian portion of the Harz mountains, aggregating 134.350 acres, of which four-fifths, or 107,480 acres, are under spruce, during the present century (up till 1870) over two millions of mature spruce were thrown by wind, or the equivalent to full crops on about 10,500 acres, nearly 8 per cent. of the total area. He also estimates that wind and snow combined have during the present century destroyed at least four millions of stems in the tree-forest stage of growth, without including those that have been merely damaged by wind or snow, and have consequently fallen victims afterwards to bark-beetles, which first of all attack the sickly stems, breed there, and then attack sound and healthy trees, unless all unsound individuals are removed.

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Pure Forests of Spruce.---Localities with a short period of vegetation being the natural home of spruce, its cultivation in pure forests in Scotland would seem advisable wherever the upper soil has the requisite moisture. In the generally damp climate of both the lowlands and highlands of Scotland with their comparatively short summer, the factors are given which hold out promise of the normal development of spruce, although its growth may perhaps not be so rapid as in warmer southern localities. At higher elevations or in the far north it takes perhaps a hundred to a hundred and twenty years to attain the same average dimensions in pure spruce forests that can be arrived at in lower or warmer localities in eighty to a hundred years, but this shortening is to a great extent counterbalanced by the better quality of the timber produced.
Tempted by the remunerative promises held out, spruce forests have often been formed on heavy loams and clays, or on sandy soils apt to suffer from want of moisture. When the climate is mild, and the period of vegetation prolonged through warm spring and autumn weather, plantations show rapid growth in youth, which, however, does not always continue throughout the whole period of rotation, but not infrequently shows signs of loss of energy about the fortieth to sixtieth year. The too rapid development during the youthful period produces soft wood of indifferent quality, which offers but little resistance to `dangers threatened by snow, by the attacks of insects, through infection with fungous disease, or to diseases originating in other causes. Although of course this is by no means necessarily the case, it is not unusual to find such spruce woods early interrupted in canopy, and unable to afford sufficient protection to the soil, so much so in fact that their clearance may be advisable before they have attained sixty years of age. On such localities spruce is not necessarily out of place, but may, grown in patches along with a ruling species for which the soil and situation are more suitable, attain very satisfactory growth, and assist very materially in increasing the ultimate returns from the crop. The periods of rotation of spruce usually vary from seventy to eighty up to a hundred or a hundred and twenty years, the former supplying the ordinary assortments of timber requisite for building purposes, the latter yielding large squares. Local demands of course to a great extent determine the most remunerative period of rotation, but where the forests are extensive, fixing the fall at an early age is apt to swamp the market with small timber, whilst entailing the harvesting of large quantities of top-ends and small material of very little value. The postponement of the fall to a hundred and twenty or a hundred and forty years, for the production of large- girthed timber, can only be advisable in very sheltered localities, owing to the dangers to which the spruce then becomes exposed; and for the same reason the retention of standards, when mature crops are being cleared, is always combined with more risk than is prudent. Here again, however, the same end can be better attained by growing the spruce in admixture with other species, such as the beech, silver fir, or pine.
The thinning out of pure forests of spruce is usually confined to the removal of suppressed individuals, and of those likely to be immediately suppressed, and in localities where damage from snow is not improbable the operations must be conducted carefully. As the natural habit of spruce is to grow in close canopy, any premature removal of poles can only be an unnecessary diminution of the number of individual stems per acre. In the weedings and clearings which take place in young crops before thinnings are begun,
(Under clearings and weedings are classifiable all the operations in young woods which necessitate an outlay that cannot be covered by sale of, the material cut out; when the costs involved are covered, or more than covered, by the proceeds obtained, the operations are then properly termed thinnings. ) all soft woods and coppice shoots of trees forming part of the former crop should be removed in order to avoid the formation of blanks later on,---birches if left standing often do great damage by rubbing and chafing leading-shoots of the spruce,---and when the crop has been formed by thick sowing or planting in wisps of two or three on the poorer classes of spruce soil, these early protective measures for improving the growth of the young stock often occasion great trouble and considerable expense. In the thickets formed by planting in wisps on inferior soil, it is especially necessary to repeat the thinnings as frequently as possible, in order to assist nature in the selection of the predominating stems to form the future crop; the longer such operations, are delayed the more difficult and expensive do they become as the roots interlace and the lower portions of the stems grow into each other. Localities exposed to the danger of snow-accumulation require the most careful thinning, and even dominated individuals should be left standing, as reserves in case of accidents, if their crowns are' still green; broken stems also should not be removed if three or four green whorls give hope of some side spray assuming the rôle of leading-shoot. Where a strong head of game is maintained, deer often do greatest damage by stripping the bark in woods directly they have been thinned, and in particular just after the first time of thinning out; from such wounds spruce suffers more serious permanent damage than Scots pine or silver fir, owing to its weaker recuperative power. For the same reason the removal of green branches with the bill or axe is not advisable; when it is desired to remove branches for the purpose of producing clean-stemmed timber free from knots, the operation should invariably be performed with the saw in the case of both living and of dead branches, and confined to those under 4" diameter. 
Soils that are somewhat inferior for the production of mixed forests of broad-leaved species, or areas where long- continued or badly managed copse, or coppice under standards, has allowed the soil to become more or less deteriorated, frequently afford good localities for the growth of spruce in pure forest, although it is not advisable to confine it to soils below the average in quality if the production of the larger, more valuable, and under certain circumstances more remunerative assortments of timber be desired. Where pure forests of spruce exist on an extensive scale over large tracts, it is exceedingly desirable to frame the working plan so that the annual fall should take place in several places apart from each other, instead of being combined in one large area annually, as it is to a great extent the latter method of procedure which causes so many of the drawbacks under which pure forests of spruce suffer. With the cessation of total clearances over large areas it is beyond all doubt that dangers from insects during the youthful period of growth, from snow during the pole-forest stage of development, and from wind when approaching maturity, would all be practically, and very considerably, lessened by the formation of several blocks, each with its growing stock of successive annual crops from one to eighty or a hundred years according to the period of rotation fixed on, in place of having the total area divided simply into eighty or a hundred compartments as the case may be, from the oldest of which a total clearance of the mature timber is annually made.
The usual method of regeneration of pure forests of spruce is, as above indicated, total clearance with artificial reproduction, except at very high altitudes where considerations of treatment are usually secondary to those relative to the general economic value of maintaining the higher mountains under forest in order to prevent landslips, and to regulate the flow of moisture through the soil and ensure the perennial feeding of the streams which have their sources there.
The unsuitability of the method of natural reproduction under parent standards that is customary in the case of the other two densely foliaged shade-bearing species, silver fir and beech, finds easy explanation in the indifferent resistance which the spruce is able to offer to the violence of storms, otherwise the diminished increment that is attained by the young crop during the earlier stage of growth would be amply compensated by the protection against various dangers which the parent trees would secure to their progeny during the first ten to twenty years. Experience has, however, shown that attempts at natural reproduction in this manner, except in very sheltered localities, usually lead to the parent standards being thrown by wind, when a rank growth of weeds soon covers the soil, and chokes a large proportion of the seedlings; and again, when the reproduction takes place by sowing or planting, the retention of standards is more likely to be injurious than beneficial.

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Mixed Forests with Spruce as the Ruling Species. ---Spruce is found forming pure forests over very extensive areas, as it is one of the species which can thrive and attain normal development without an admixture of other kinds of timber trees in the crop. At the same time there is hardly any other species of forest tree in Britain which gains so much as the spruce by the formation of mixed forests, both as regards the unquestionable, protection thus afforded to it against dangers, whether of organic or inorganic nature, and in respect to the stimulus thereby secured for the total production of timber per acre, and the better quality of the timber produced.
In its Alpine home, the larch is frequently to be found naturally associated with spruce at the higher elevations, although artificial admixture of these two species in other localities has often been far from satisfactory. Throughout the mountainous tracts of eastern France, and of central and southern Germany, in particular in the Black Forest, mixed forests of spruce and silver fir are a favourite form of timber crop,---except on the Harz mountains, where the climatic factors do not seem to be favourable to the development of the latter. Beech is also often an associate of the spruce, and is to be found frequently in mixed forests of spruce and silver fir. Towards its northern and `eastern limits, the species chiefly found growing along with spruce is undoubtedly the Scots pine. These are the trees which are usually found growing as subordinate species in mixed forests where spruce forms the ruling species or matrix, and although other mixtures have been tried artificially, the above-named are those which hold out the best sylvicultural and economical promises.
When the silver fir finds the soil and situation congenial, it is the most important associate of the spruce; and though during the earlier years of growth it must be granted some protection against the more rapidly developing spruce, it requires no special tending throughout the later stages of growth. In many respects it adapts itself better for admixture with the spruce than with any other ruling species of forest tree. In tree-form and natural development they have many close resemblances, but as silver fir is deeper-rooted than spruce, the two species, admixed, can develop unhindered a much larger aggregate of roots than if either species were grown in pure crop; and as this maximum of root-system has practically the opportunity of drawing the requisite supplies of nutriment from two different layers of soil, it follows naturally that the number of stems and the total production of timber per acre will under ordinary circumstances be considerably greater than can be shown by pure forests of either species. By interlacing of the two root-systems also, no inconsiderable support is given to the shallow-rooting spruce against windfall.
The main condition for the formation of mixed forests of spruce and silver fir is a good deep soil without excess of soil-moisture, as unless that essential condition be satisfied the latter is unable to maintain itself against the former. Even although developing slowly at first as compared with Scots pine, spruce has a more rapid early growth than the silver fir, and the advantage thus won it maintains throughout the pole-forest and into the tree-forest stage of development. Although the silver fir can thrive in the side-shade cast by the spruce, measures must be taken to prevent the latter shooting so far ahead as to form canopy above the silver fir in the thicket stage of growth, otherwise the latter dies off. Where the two species are planted out alternately in equal numbers, the silver fir soon gets defeated in the life-struggle, unless the soil is of better than average quality. Even when the admixture takes place by planting in alternate rows, the silver fir transplants should have the advantage of being older than those of the spruce. Where such mixed forests are formed or reproduced, it L is advisable to allow the silver fir the advantage of five to ten years of growth in order to enable it to protect itself against the spruce without necessitating considerable outlay for tending,
Beech is not of so much importance as silver fir as a minor  species in spruce forests, for though its root-system is heart-shaped like that of the former, its general habit of growth and development as a forest tree is greatly different from that of the latter; it requires to be grown in groups or patches in order to maintain itself at all against the much quicker growing spruce. Although it yields better fuel than any other species of forest tree, the wood of the beech is in poor demand as timber for technical purposes, so that in Britain it will usually only be found in forests on account of its soil-improving qualities. 
Important though the considerations regarding increased annual production and better quality of timber be, yet the chief advantages to be gained through the introduction of silver fir and beech are beyond all question or doubt the greater security afforded to the spruce in respect to all the dangers and enemies to which this species is exposed. In localities where spruce can thrive safely till maturity, measures for increasing the production are hardly of the first importance, as the returns from pure forests of spruce are in themselves so good that further outlay for the introduction of a minor species might often seem uncalled for; but where, as in most localities under spruce, storm, snow and ice-accumulations, attacks of insects, fungous diseases especially (Trarnetes radiciperda and Agaricus melleus), and other dangers cannot be left out of reckoning, an intermixture of one or other of these species - and on suitable soils and situations preferably the silver fir, - is in the highest degree advisable in the light of recent experience throughout Germany.
On many parts of the northern slopes of the Bavarian Alps, the larch also occurs as a subordinate species along with silver fir and beech in spruce forests. Its growth at first is much more rapid than that of the spruce, and on deep fresh soil the advantage thus early won is maintained till the sixtieth to seventieth year, or under favourable circumstances longer, although only too often it is caught up and overtopped by the spruce. Where the soil, however, is wanting in depth and strength, or where, as in Britain, both species are removed far from their natural homes, and cultivated under conditions in many respects dissimilar from their normal requirements, it not infrequently happens that the spruce, stimulated to lively growth in height, catches up the larch as early as the twentieth to thirtieth year, when nothing remains but to cut out the latter, and allow the spruce to form pure forest. Even when the larch has been introduced in patches among the spruce, little can be done in such cases to protect it, as its further growth is prejudiced by the side-shade, and it can no longer develop satisfactorily. As a rule, the larch should only be grown in spruce forests on parts where the soil is of better quality than the surrounding ground, and on such patches it generally thrives better in groups than when planted out alternately with spruce, or only simply here and there,---although it maybe remarked here that Burckhardt recommends its being planted out singly only, except along the edges of compartments where it may form rows or belts. The groups should not, however, be too large, as otherwise there is the same tendency towards crooked, sabre-like growth as is. characteristic of pure larch forest away from its Alpine home. Experience shows that when grown along with spruce, the larch is less liable to be attacked by fungous disease (Peziza Wilkommii) than when it forms pure forest. As a rule large transplants should be used in introducing the larch into spruce woods, so as to assist in giving it the greatest possible advantage in growth, - but unfortunately where roe-deer are maintained, these are specially sought out by the bucks at the. time of brushing the velvet from the horns in early summer, and much damage may be caused in this way.
Scots pine is seldom to be found as a minor species on the better classes of spruce soil, where the preference is usually given to those others already mentioned; but it forms a valuable associate on the poorer qualities of soil, and wherever the satisfactory development of the spruce is likely to prove questionable. The rôle that it then plays is partly that of a purely subordinate species, partly that of a nurse or protector. In such cases the object in view is to raise the spruce in as large a quantity as possible, but at the same time to have the pine represented to as great an extent as can be grown along with the spruce, or as is necessary for the maintenance of closed forest. That, under such circumstances, what was originally intended as a spruce forest with the admixture of Scots pine, ultimately approaches maturity as a crop of pine with admixture of spruce, can easily be understood, as on such debatable land consideration must be duly given to the factors influencing the growth of both species at many critical periods of the life-history of the growing-stock. Any stencil-like regularity and uniformity of treatment of such mixed crops is out of the question, and it can only be expected that, with proper and prudent treatment, the mature fall will consist here and there of spruce with pine, and in other parts of pine with spruce intermixed. When there is doubt about the soil suiting the spruce, it is perhaps a good rule always to form the young crop by means of an equal admixture of both species in rows or bands proceeding later on with the clearings and thinnings as may seem advisable each time these operations are under consideration.
Such cases of doubtful success are just as frequent on the dry slopes of low hills and uplands, as on the more level tracts or plateaux. Pure pine forests are often just as much out of the question as pure forests of spruce, for whilst the latter is slow in closing up to form canopy, and inactive in growth generally, the former is apt to become interrupted in canopy at too early a stage of development, and to fail in affording to the ground the protection so specially requisite on soils of inferior quality. In mixed forests consisting equally of spruce and Scots pine, - or of spruce to half the number, and Scots, black and Weymouth pines forming the other half, - the best possible attainable results are perhaps achievable, the ultimate tending of the crop being dependent on the relative development of the different species. In Hanover it was usual, at the time when sowing stood in greater favour among sylviculturists than planting, to mix and sow spruce and Scots pine seed in the proportion of five to one, the pines being regarded [solely as nurses, and cut out as soon as they began to inconvenience the spruce with their shade.
That, in equally mixed spruce and pine forest, the latter often becomes the dominant species, is due to inefficient tending more than anything else, for, unless some special attention be paid to the spruce during the clearings and thinnings, it either remains dwarfed as underwood, or at any rate has no fair chance of developing until the canopy of the pine becomes naturally interrupted. Many mixed woods of this description yield good returns if the pines are cleared away wherever the spruce shows need of freer enjoyment of light and air, and stems are left only here and there on the better patches to increase rapidly in girth over the well-protected soil.
The oak is not naturally a common associate with the spruce, from which it differs essentially in many sylvicultural characteristics. The oak is generally found on the milder situations on plains and uplands with a long warm period of vegetation, the spruce at higher elevations and on shallower soils. Still, in many parts of northern Germany such mixed forests do exist, and are often spoken well of. Even when oaks are given a few years' advantage at first, they are soon overtaken and topped in growth by the spruce, unless they are planted in clumps of considerable diameter. If planted in rows or small groups, though they may reach the pole-forest stage of growth evidently thriving and well above the spruce, they seldom maintain these advantages till maturity, but have usually to be cut out long before they attain good marketable dimensions.
Softwoods are often found associated with spruce, as nurses where the reproduction of the latter is difficult, or as protective standards in situations exposed to frost. But their artificial production is seldom necessary, as they usually occur self-sown, and if not, a more desirable substitute for parent shelter can generally be found in the pine. More frequently they in reality become weeds, whose coppice-shoots occasion much trouble and annoyance. This is particularly the case with coppice-growth of the birch, whose long whip-like twigs damage the leading-shoots of the young spruce growing around; but where seedlings of birch occur merely scattered here and there individually throughout spruce woods in places where late and early frosts are to be feared, their retention till they are caught up in growth by the spruce, often yields good preliminary returns as well as useful aid sylviculturally.

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Formation and Reproduction of Spruce Forests. ---Except at high elevations, where the ordinary methods of reproduction of spruce forests cannot be carried out~ and where their profitable working must be subordinated to the main object of maintaining the mountain-tops under woodland, there are three distinct forms of reproduction, all of which are practised in regular annual falls. These are:
    1. Natural reproduction under parent standards.
    2.Total clearance in narrow strips, with natural reproduction from neighbouring woods.
    3.Total clearance of annual fall, with artificial reproduction (usually by planting).
Natural reproduction under parent standards is especially practicable in respect to mixed crops of spruce with silver fir and beech, but is, however, also adopted in pure spruce forests on level soil, where late frosts or attacks of cockchafer grubs (Melolontha vulgaris) are to be feared on an extensive scale, and experience has further shown that in forests thus reproduced the dangers from. Curculionidae are likewise diminished. Other local circumstances must of course be taken into consideration, and this method of reproduction will often recommend itself in outlying and sheltered localities, where the proprietor does not wish to incur the usually moderate costs of artificial regeneration. The results of natural reproduction under parent standards are varying. In some situations the parent standards are not much exposed to the violence of storms, but in most localities this is unfortunately not the case. The young crop often varies much in quality; in some situations a moist soil is favourable to germination and the seedlings stand too thick, whilst in other places reproduction is slow and unequal, resulting in thin patches of seedling growth of different ages, necessitating some artificial assistance, and adding considerably to the costs of tending later on.
The method of total clearance with natural reproduction from neighbouring woods, was formerly much more frequently adopted than is now the case. The fall for reproduction should not be more than 100---120 yards broad, and must of course be so located that the adjoining mature woods lie to the windward, in order that seed may be evenly shed over the area when the cones open with dry warm winds in late spring and early summer. Some measure of soil preparation for the reception of the seed is absolutely requisite, and when seed-years turn out disappointing, artificial reproduction or assistance becomes a necessity to a greater or less extent, as otherwise a rank growth of grasses and other weeds covers the soil and shuts out the hope of seedlings being subsequently able to force their way through these successfully.
The total clearance of the annual fall of the mature crop with artificial reproduction by planting, is now the usual method of treatment of spruce forests in Germany. Independent of seed-years, untrammelled by considerations regarding the protection of standard parent trees against the violence of storms, and far less threatened with danger from growth of weeds, reproduction can thus be carried out quickly and satisfactorily at a moderate cost, whilst the extraction of the mature timber is easier, and the grubbing up of the roots less difficult in localities where there is any good market for fuel. This method has many advantages to recommend its adoption,---it is an easy system, involving the minimum outlay for supervision, tending, and ultimate harvesting of the crop, the annual fall of timber is regular, and varies little in quality or cubic contents, and the working plan is based on the simplest and safest of all foundations, viz, equality of the areas (modified according to their relative productive capacity) from which the mature crop is annually cleared; it has, however, drawbacks and disadvantages which have previously been referred to.
In whatever manner spruce forests are reproduced, a general principle should be followed of not making the annual fall comprise too large an area, as it is in every way of unquestionable advantage to have a series of self-contained blocks, each comprising within itself crops varying from one to eighty or a hundred years, instead of one large block simply divided into eighty or a hundred annual compartments or falls.

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Natural Reproduction.---When reproduced naturally under parent standards, no preparatory fellings are necessary to stimulate the production of seed and prepare the soil for its reception. When a good seed-year seems favourable for reproductive fellings, they are made so as rather to resemble those in beech and silver fir, than in Scots pine forests; but on account of the danger from wind the number of trees left per acre is greater, only from ¼ to of the total number of trees forming close canopy being removed, so that during storms the crowns can afford each other some measure of support. On moist soil, a lighter disposal of the parent trees would also favour a rank growth of weeds, which is more prejudicial to young spruce than even a considerable degree of shade from lofty standards. The period of reproduction is much shorter than with beech or silver fir, as the seed-years are more frequent, and the amount of seed produced greater, besides which the young seedling growth is not so absolutely shade-demanding as with these other species. From the pine it also differs essentially, not only in the more abundant, though not more frequent, production of seed, but also in that the seed ripens in about six months, in place of being delayed till eighteen months after the flowering.
The shape that it is advisable to give the area to be reproduced is dependent on the extent of the danger from wind; the greater the danger, the more should reproduction take place in long narrow strips on which the number of trees along the middle should be greater than towards the edges, so as to ensure speedier regeneration and earlier clearance of the parent trees, in order to minimise the damage caused to the ~ seedling growth at the time of extraction. Where good patches of self-sown spruce occur they should be retained, but all other species of trees should be cut out. When practicable, large branches should be sawn off to decrease the leverage obtainable by the wind, but care should be taken to carry out this operation during winter, in order to prevent the outflow 6f sap from the wounds. Whatever soil-preparation can be conveniently undertaken yields its reward in easier and better growth of seedlings ; the layer of thick moss should at any rate be removed with the rake. In Prussia, breaking up of the soil roughly into clods in bands or strips one to one and a half feet broad and six feet apart has been found a judicious outlay, the operation being performed in the autumn of the seed-year.
The clearance of the standard parent trees commences in the winter of the year following the seed-shedding, and the extraction should take place, so far as possible, whilst snow lies on the ground, in order to minimise the injury done to the seedling crop; clearance must be effected as speedily as possible, as the danger from wind increases greatly when once this operation has been begun; even in sheltered localities the final clearance should be completed by the time the seedlings have attained a height of one foot. Where reproduction has not been equally and uniformly successful, it is not advisable to retain the standards; blanks can easily be filled up by sowing or planting, or an excellent opportunity is thus given for introducing other species such as silver fir, beech, pine or larch, whose admixture along with spruce has been shown by experience to be so desirable for many good reasons. Douglas fir should also yield good results.

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Artificial Reproduction and Formation.---During the last century, sowing was the usual method of forming or reproducing spruce forests artificially, but towards the beginning of the present century planting, in place of being confined merely to the filling up of blanks in sowings, became a rival of the older method, and for the last fifty years it has been the favourite system. Sowing, besides not always being so successful, is on the whole not so very much cheaper than planting that one can afford to overlook the difference of two to four years' growth won for the future crop when seedlings or transplants are utilised. But planting of spruce has other advantages over sowing. Plantations suffer less from rank growth of grass, run less risk of being lifted out of the ground by frost, and also suffer less where large herds of deer are maintained, whilst they can be opened earlier to grazing; they thrive as a rule better than young crops raised from seed, and can be formed in autumn as well as in spring. 
The material for filling up the blanks in crops raised from seed was formerly usually taken from reserve plots or temporary nurseries prepared by sowing thickly in the proportion of about 150 lbs. of seed per acre actually sown. By the time the seedlings were four or five years old they could only be used in wisps of three to five, in place of individually, as they had grown quite entangled, a method that is even now intentionally practised under certain circumstances, but which is hardly recommendable, as it renders subsequent tending difficult, and interferes too often with the normal development and the early selection of predominating poles.
The distances at which seedlings and transplants were planted out in the great home of the spruce, the Harz mountains of central Germany, has varied at different times. At first plantations used to be made at 2½ feet x 2½ feet, but later on the plants were made to stand at from 3 feet x 3 feet to 5 feet x 5 feet, the wider distances being preferred where there was heavy snowfall; recent experience in Germany has shown, however, that 4 feet x 4 feet is preferable to any wider distance.

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Sowing. ---Although planting is now generally admitted to be preferable to sowing, yet under certain circumstances the latter method finds its proper uses, as, for example, where seedlings or transplants are not conveniently obtainable, or where the soil is too rocky or otherwise unsuitable for planting, or when a good market for small material like pea-sticks can be tapped early and remuneratively. More seed per acre is used for spruce than for Scots pine, for not only do fewer seeds go to the lb. (55,000 to 57,000), but a denser crop is also desirable. As a rule about 10  to 12 lbs. per acre are used, although this quantity must be increased in proportion to the magnitude of the danger from drought, weeds, or lifting of the seedlings by frost. Sowings are less frequently made broadcast than in rows, or strips, or on small patches, and clean seed is now alone used, a soil-covering not exceeding a quarter of an inch being provided by light raking. The operation is carried out towards the end of April or the beginning of May, as a rule, and only exceptionally in autumn. Except where rank growth of weeds demands broader strips the soil is generally prepared
to a breadth of 1½ to 2 feet and in rows 3 to 4 or at most 5 feet apart; patches are usually 1½ to 2 feet square and about 4 feet apart. On hilly situations the rows should run horizontally, and not vertically, to prevent the seed being washed away.

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Planting. ---The best results are obtained with transplants from regular nurseries, although younger seedlings taken from seeds beds, or wisps (three to five) from rills in temporary nurseries, also at times find favour. Plants from two to five years are usually put out, in preference to yearling seedlings, as the planting of the latter often costs nearly as much as if two-year-old plants be used, whilst the results are generally not so good as with these.
Notching, and the use of naked seedling or wisps, are only suited for the more favourable soils of a light character; on tenacious soils, or where there is a strong tendency to growth of weeds, this otherwise cheap and good method is not advisable. In general, too, this method seems to affect the rootlets of the spruce to a greater extent than those of the pine, which latter do not so long retain the flat shape induced by the pressure employed at the time of planting.
The use of transplants with earth attached is on the whole far more advisable, and usually leads to better results, than the use of naked seedlings; and in localities where growth of weeds does not call for the use of older material, the operations, of transport and planting can be carried out at a very reasonable rate if the preparation of the holes on the area to be planted, and the lifting of the material from the nurseries, be carried out with small cylindrical spades (Heyer's -
These useful instruments were first, described in a report "On the Corsican Fir,' printed in the Transactions of the Highland and Agricultural Society, 1876),  which besides have the additional advantage of rendering too deep planting impossible,---one of the worst, but most common mistakes made in regard to the spruce, particularly in tenacious soil.
Planting should take place in spring to as great an extent as is practicable; but at high latitudes or elevations, or where a moist soil has first to get rid of some of its superfluous moisture, autumn planting also yields good results. On windy situations, and on dry soil, the most favourable time for planting is is spring, just before the buds flush and form the new shoots. Plantations formed in autumn are less able to resist the action of frost during their first winter than those formed in spring, whilst the plants can more easily be pulled out of the ground by deer.
The best average distance between the plants has been found to be four feet; whether greater or less distances recommend themselves in any particular case depends upon local circumstances and on the funds available, the principal advantages of closer planting being in the speedier attainment of close canopy and protection of the soil, in greater freedom from branches, and in earlier returns from small material removed during the operations of thinning out. When requisite, it is better to give the individual poles more growing-space by means of the bill or the axe, than to endeavour to secure it from the very outset by means of planting at wide distances; but where there is a poor market for pea-sticks and the like, and where labour is neither plentiful nor cheap, a preference will often be justified in favour of somewhat wider planting. Where the soil is fresh and good, and only sound four to five-year- old transplants are set out, wide planting at 5 feet x 5 feet should be sufficient; but care must be taken to fill up any blanks promptly, as with only 1,742 plants per acre to start with one cannot afford to lose any prematurely (vide tables on pages 22 and 43). But where, on the other hand, the soil is dry, or where rank growth of whortleberry, heather, or other weeds has to be contended with, considerations as to the speedy formation of close canopy and suppression of weeds may determine in favour of closer planting, for 2,725 plants per acre at 4 feet X 4 feet effect the purpose much more quickly and effectually than 1,742 at 5 feet x 5 feet. On dry slopes, or deteriorated soil, even closer planting is advisable when funds are at disposal, although a judicious admixture of Scots pine along with the spruce often leads to the attainment of the object in view at less cost than close planting of spruce alone. When planted for shelter, as a protective mantle along the edge of pine or other forests, they should not be put out closer than five or ~ix feet in order that the individual trees may develop fully in foliage and be able to maintain themselves against storm-winds by a free formation of their' root-systems towards the windward side.
Whether the setting-out of the plants should take place in squares, or triangles, or rows, is of less importance than the actual number of plants per acre; practically, planting in squares is the usual method adopted, although in situations where plantations are liable to suffer from accumulations of snow or ice, planting in rows of 6 feet x 4 feet, or 6 feet x 3 feet are said to yield the most satisfactory results. Close planting increases the danger from snow, whilst ice causes most breakage when the individual plants have a fair amount of free growing-space.
As previously remarked, notching is not so well applicable to the spruce as to Scots pine, but recommends itself on account of its cheapness wherever the nature of the soil is suitable. The usual methods adopted are planting by means of Heyer's cylindrical spade for one and two- year-old seedlings and three-year-old transplants, and pit-planting,---or on wet soil, tumping or planting on. mounds --- when older material is put out.

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